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This is an open access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Vertical discontinuities were detected in several panels in this article, including panels of Figs 3A, 3B, 3C, 4B, 4F, 4G, 5B and S1B. PLoS Biol 18(10): e3000972.

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